This is usually for high power V8s. Like Mick said, it's an EPA thing.
My 5.0 has a 1-4 "skip shift" feature. If you aren't accelerating quickly the transmission locks out 2nd gear and suggests a 1-4 shift. Doesn't really bother me much because I usually shift 1-3-5-6 or 1-4-6 anyways when I'm just driving normally. It is easily removed with a handheld tuner if it bothers you.
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- Steve
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