The difference in sound from an iPod v. Zune comes down to the internal parts. The DAC in the Zune is better, they have both been torn down and proven, but unless you carry around a portable amp with nice headphones you will never notice the difference.
Now in a car, with a good set of speakers, amp, etc., you might tell a difference but the sound stage would vary from car to car. Also it is relative to how your music is encoded, MP3 v. WMA v. FLAC v. AAC.
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